https://bmcdevbiol.biomedcentral.com/articles/10.1186/1471-213X-6-41
https://bmcdevbiol.biomedcentral.com/articles/10.1186/1471-213X-6-39
https://bmcdevbiol.biomedcentral.com/articles/10.1186/1471-213X-6-40
https://bmcdevbiol.biomedcentral.com/articles/10.1186/1471-213X-6-38